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Spring has come and the ponies of Ponyville are getting excited because it is time to go to Wisteria's garden party to discuss plans for the Spring Promenade. Spring Promenade is one of the biggest parades of the whole year and it requires a lot of gardening to produce all the flowers needed to decorate the floats for the parade. With help from the flying “Breezies,” Tiddlywink, Tra La La and Zipzee, the gardening begins in earnest. Seeds are planted, watered and cared for and soon, flowers begin to bloom all around Ponyville. One day while tending the garden, Wisteria discovers a weed. When she can't pull it out, she asks Pinky-Pie to help her. The two ponies dig and dig until the hole becomes so large that they fall down into it. Zipzee follows along to help them and the three friends find themselves in a dark underground passageway where they follow a bad smell to the site of a beautiful, rare, blooming flower. Living in the dirt under the beautiful flower, they find Spike the dragon and realize that the bad smell is coming from Spike who informs them that he has been sleeping there for over a thousand years. He goes on to tell him that according to the legend, he would be awakened when someone touched the rare flowerand then that someone would become a princess. It was Wisteria who touched the flower and thus she is proclaimed to be a princess. When the ponies and Spike return from the underground to Ponyville, everyone is thrilled to learn that Wisteria has become a princess and they rename the parade “Princess Promenade” in her honor. Spike begins his task of guiding and protecting the new princess. Under Spike's guidance, Wisteria learns that she can't help with the preparations for the parade, she can't dig in the garden and she can't even leave the castle. In the meantime, the ponies are struggling to get things ready for the promenade but they miss Wisteria's knowledge and help. Wisteria's unhappiness with all the restrictions causes her to say, “I want to be me again.” Spike cannot believe his ears. He says that everyone want to be a princess or a prince. He asks how she could possibly quit. Wisteria tells him she has figured out the way to stay a princess and “still be me.” On the day of the Promenade, Wisteria stops the parade to make some important announcements. She shares the important lessons that she has learned from her time as a princess and she announces that she is making everyone in Ponyville a princess. Everyone celebrates the beautiful Princess Promenade.
The film is moved along by several songs and features the classic pinks, purples, greens and other soft but lush colors that are part of the My Little Pony mystique. Some parents may conclude that the amount of primping, e.g. hair-dos, manicures, pedicures, and dressing-up, is overdone and sends a message to girls that these behaviors matter too much. Nevertheless, the message about being true to yourself is the most important message and it comes through loud and clear.
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